Table 4 in New record and new species of Laubierpholoe Pettibone, 1992 (Annelida, Sigalionidae) from the soft bottom of submarine caves near Marseille (Mediterranean Sea) with discussion on phylogeny and ecology of the genus

Table 4 (continued on next page). Comparison of species of Laubierpholoe Pettibone, 1992.L. antipoda (Hartman, 1967)L. indooceanica Westheide, 2001L. maryae Pettibone, 1992L. riseri Pettibone, 1992L. swedmarki (Laubier, 1975)L. massiliana Zhadan sp. nov.Type locality, ecology, distributionSouth Atlantic Ocean, off South America, Tierra del Fuego, in 57–494 m; off New Zealand, in 86–95 mSouth west Indian Ocean, Seychelles, Mahé, reef flat, South India and the Seychelles; extremely clean coarse coral sandNew Zealand, South Island, Kaikoura Harbor, Rubies Reef, in 12 m, coarse broken shell; Aquarium Point, Otago Harbor, intertidalNew Zealand, North Island, Leigh, middle of Goat Island, coarse broken shell in Amphioxus sand, in 20 mNortheast Atlantic Ocean, Castle Island, Bermuda, in 2–8 m; CubaMediterranean Sea, Gulf of Lion, the Calanques near Marseille, marine caves, silty and sandy sedimentBody length, mm1–30.6–0.8752.8–31.5–1.81.60.8–1.2Number of segments13–2713–1527–2919–262616–19Prostomium and anterior hornsoval, bilobedbilobedoval, bilobed, with anterior lobes projecting anteriorlyoval, bilobed, anterior lobes extended anteriorlyrounded, bilobed, with lateral hornsoval to trapezoidal, bilobed; anterior horns arising medially from ventral tentacular cirriMedian antennastyle enlarged basally, about same length as dorsal tentacular cirristyle with a distinctly narrower distal part, considerably shorter than dorsal tentacular cirriwith style about as long as prostomium, shorter than dorsal tentacular cirrismall ceratophore, style short, with filamentous tip, shorter than dorsal tentacular cirrismall ceratophore and short style, twice as short as dorsal tentacular cirrilarge ceratophore and short style, shorter than dorsal tentacular cirriTentacular cirridorsal cirrus much longer than ventraldorsal cirrus much longer than ventraldorsal cirrus much longer than ventraldorsal cirrus much longer than ventraldorsal cirrus much longer than ventraldorsal cirrus slightly longer than ventral or cirri of about same lengthEyes2 pairs, large, separate2 pairs, lensed, of nearly identical sizeabsent2 pairs, large, anterior pair larger2 pairs, anterior pair larger2 pairs, very closely situated, anterior pair largerElytrasquarish to subreniform, smooth, with up to 7 papillae along lateral margin, sometimes few papillae on surfaceirregularly oval, covering trunk and parapodia, with 3 to 4 short, capitate papillae along anterior margin and 3 to 5 on surfacelarge, oval, covering dorsum, delicate, with few papillae on lateral border and few on surfacelarge, oval, covering dorsum, with few lateral papillae; elytra on posterior segment small, enclosing pygidiumoval, not covering middorsum; transparent, with 5–8 papillae on external border and few on surfacelarge, round, transparent, with few oval to cirriform papillae along lateral and posterior borders and on surfaceNotopodiasubconical acicular lobe, smaller and shorter than neuropodiaconical acicular lobe; shorter and smaller than neuropodiaconical acicular lobe, about as long as neuropodiumsubconical acicular lobe, longer than lobe of neuropodiumsmall, conical acicular lobe on anterodorsal side of larger neuropodiaconical acicular lobe; shorter and smaller than neuropodiaNeuropodialong conical prechaetal acicular lobe and short rounded postchaetal lobeconical lobeconical acicular lobesubconical acicular lobeprechaetal conical acicular lobe and shorter rounded postchaetal lobeconical acicular lobeNotochaetaenumerous, slender, slightly curved, with capillary tips; upper ones shorter, more strongly curved, ventral longerfew (4–8), exclusively slender, slightly curved, with minute serrations along convex edgenumerous, slender, capillary, slightly curvednumerous, slender, capillaryfew (2–4), forming fan-shaped bundle, slender, with series of small denticlesfew (3–6), slender, capillary, straight or slightly curved, from smooth to finely serratedNeurochaetaeshafts with row of subdistal denticles; blades short, with row of minute denticles and entire, falcate tips; upper ones with blades slightly longer than lower onesshafts distally minutely spinose; in CH1 all blades minutely serrated; from CH2 two uppermost supraacicular chaetae with blades long, serrated; other supraacicular ones with blades long, smooth, sub- acicular ones with blades short, smooth; hook-like blades in five posteriormost segmentsshafts without distal spines; few supraacicular chaetae with blades long, minutely spinose and blunt tip, other supraacicular ones shorter, coarsely serrated, tapering to sharp tip; subacicular with blades shorter, falcate, minutely spinoseshafts without distal spines; supraacicular chaetae with blades long, tapering to capillary tips; subacicular ones with blades short, falcateshafts with numerous distal and subdistal spines; blades short and falcate, with entire tips, upper ones longer, with more numerous spines, lower ones shorter with fewer, minute spinesshafts with numerous subdistal and long distal spines; upper blades slightly longer than lower ones; blades serrated, straight, falcate and of 2 types: supraacicular ones unidentate, subacicular ones uni- and bidentate